 The study evaluates the effect of climate factors on the reproductive phenology and success of gray wolves in North America, finding that while spring onset shifted earlier due to warmer temperatures, denning dates did not change and reproductive success was not affected by timing of denning or synchrony with spring onset. The study highlights a disconnect between climate factors affecting phenology and those affecting demography, suggesting that carnivores may be resilient to shifts in seasonality but sensitive to weather conditions affecting their prey at both local and regional scales.