 The Hippocampus is capable of encoding individual episodes and extracting regularities across experiences, which is central to category learning. Using a neural network model that mirrors the anatomy of the Hippocampus, we found that the trisynaptic pathway within the Hippocampus was critical for remembering exemplar specific information, while the monosynaptic pathway specialized in detecting the regularities that define category structure across exemplars. This article was authored by Yelena Suavey and Anna C. Shapiro.