 This study measures the relationship between stomatal traits and environmental drivers across plant communities in China, demonstrating a trade-off between stomatal density and size at the community level, with mean and variance of stomatal density mainly associated with precipitation, while that of stomatal size is mainly associated with temperature. The skewness and kurtosis of stomatal traits are less related to climatic and soil variables, and beyond mean climate variables, stomatal trait moments also vary with climatic seasonality and extreme conditions. This article was authored by Tseng Tseng Lu, Lauren Sack, Ying Li, and others.