 Holocene peat model determines vegetation community dynamics and annual net primary productivity based on peat depth, as a proxy for nutrients and acidity, and water table depth. Annual peat, carbon accumulation is the net balance above, and below ground productivity and litter slash peat decomposition. Vascular plant functional types accounted for 65% of total NPP over 8,500 years, but only 35% of the final, contemporary, peat mass. This article was authored by S. Froking, N. T. Rulet, E. Tutila, and others.