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It's a marxophone. It's a zither with courses of two strings like a mandolin. You press a key, and a hammer bounces on the strings, giving a tremolo effect.
There is mandolin throughout the song, and the autoharp starts about 4:29.
I'd say the intrument is a zither- I played one as a teenager and while that was a while ago as I recall it sounded exactly like that. An autoharp is another likely candidate.
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There is mandolin throughout the song, and the autoharp starts about 4:29.