Uploaded by TutorVista on May 3, 2010
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leaf is an above-ground plant organ specialized for photosynthesis. For this purpose, a leaf is typically flat (laminar) and thin. There is continued debate about whether the flatness of leaves evolved to expose the chloroplasts to more light or to increase the absorption of carbon dioxide. In either case, the adaption was made at the expense of water loss. In the Devonian period, when carbon dioxide concentration was at several times its present value, plants did not have leaves or flat stems. Many bryophytes have flat, photosynthetic organs, but these are not true leaves. Neither are the microphylls of lycophytes. The leaves of ferns, gymnosperms, and angiosperms are variously referred to as macrophyll, megaphylls, or euphylls.
Leaves are also the sites in most plants where transpiration and guttation take place. Leaves can store food and water, and are modified in some plants for other purposes. The comparable structures of ferns are correctly referred to as fronds. Furthermore, leaves are prominent in the human diet as leaf vegetables.
A structurally complete leaf of an angiosperm consists of a petiole (leaf stem), a lamina (leaf blade), and stipules (small processes located to either side of the base of the petiole). The petiole attaches to the stem at a point called the "leaf axil." Not every species produces leaves with all of the aforementioned structural components. In certain species, paired stipules are not obvious or are absent altogether. A petiole may be absent, or the blade may not be laminar (flattened). The tremendous variety shown in leaf structure (anatomy) from species to species is presented in detail below under Leaf morphology. Periodically (i.e. seasonally, during the autumn), deciduous trees shed their leaves. These leaves then decompose into the soil
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