I'm studying wolf behaviour at the moment for an ethogram, and they dont howl when theyre sad. Theyre probably howling to strengthen bonds within the pack. They'd be whining and whimpering if they were sad :)
This video was actually really helpful. People would not think it was, but I needed to see this for a degree assignment on behaviour, and you cannot always rely on videos or seeing a dog do it...
I'm studying wolf behaviour at the moment for an ethogram, and they dont howl when theyre sad. Theyre probably howling to strengthen bonds within the pack. They'd be whining and whimpering if they were sad :)